They took the lead when Victor Wanyama rose majestically to head past Victor Valdes from a corner on 21 minutes.
Barcelona pressed for an equaliser, but teenager Tony Watt came off the bench to give Celtic a cushion in the closing minutes.
Lionel Messi tapped home in injury time, but Celtic held on to inflict Barca's first defeat of the season.
It was a momentous victory for manager Neil Lennon, who was part of the team that knocked Barca out of the Uefa Cup in 2004, and his men - banishing memories of the injury-time defeat in Barcelona a fortnight ago.
And it leaves Celtic second behind the Catalans in Group G on seven points, three ahead of next opponents Benfica and four better off than Spartak Moscow, who visit Glasgow on the final matchday.
Celtic Park was rocking at a decibel level previously reserved for the most meaningful of Old Firm derbies, but predictably the unflappable maestros of Barcelona, twice champions of Europe in the last four years, paid the partisan crowd no heed as they quickly settled into their fluid passing game.Celtic, who last featured in the group phase in 2008, made Valdes work from another corner, though Mikael Lustig's header lacked the power to really test him.
Forster, sublime in Camp Nou, reproduced that form on the hour mark as he got down quickly to push away Alexis's well-struck shot from 10 yards and the keeper was promptly back to his feet to deny the Chilean on the rebound as well.
Marc Bartra almost gifted Celtic a second goal, waiting on Valdes to come and deal with a cross ball. When the keeper chose to stay on his line, Mulgrew looked poised to volley past him, but Bartra recovered to block the shot.
The Celtic fans bayed for a red card for Song midway through the half for another foul on Miku, but the referee again let him off.
The keeper dispatched the ball upfield, Xavi failed to cut it out and Watt showed composure beyond his years to bury it past Valdes.
Celtic fans were whistling at deafening levels for referee Bjorn Kuipers to end the game, perhaps mindful of losing a 2-0 lead in the final few minutes at Dundee United on Sunday.
And Messi ensured a nervy final few minutes as he scored at the start of injury time, following up after Forster had saved superbly again, but Tito Vilanova's side could not find another goal and Celtic and their supporters were left to celebrate long into the night.
They quickly resumed their relentless probing of the Celtic defence, almost finding an equaliser as Messi played a one-two with Andres Iniesta before rifling a shot off the top of the crossbar.
The frame of the goal came to Celtic's rescue again as Alexis got on the end of a Dani Alves cross only to see his header come back off the post.
And then Kelvin Wilson scrambled the ball off the line as an Alba cross caused hearts to flutter in the stands.
Nonetheless, unlike in the previous meeting, Celtic held onto their lead until half-time and it was fully 11 minutes into the second period before Forster was meaningfully called into action, diving to his right to stop a curling Messi effort.
Forster, sublime in Camp Nou, reproduced that form on the hour mark as he got down quickly to push away Alexis's well-struck shot from 10 yards and the keeper was promptly back to his feet to deny the Chilean on the rebound as well.
Marc Bartra almost gifted Celtic a second goal, waiting on Valdes to come and deal with a cross ball. When the keeper chose to stay on his line, Mulgrew looked poised to volley past him, but Bartra recovered to block the shot.
The Celtic fans bayed for a red card for Song midway through the half for another foul on Miku, but the referee again let him off.
The keeper dispatched the ball upfield, Xavi failed to cut it out and Watt showed composure beyond his years to bury it past Valdes.
Celtic fans were whistling at deafening levels for referee Bjorn Kuipers to end the game, perhaps mindful of losing a 2-0 lead in the final few minutes at Dundee United on Sunday.
And Messi ensured a nervy final few minutes as he scored at the start of injury time, following up after Forster had saved superbly again, but Tito Vilanova's side could not find another goal and Celtic and their supporters were left to celebrate long into the night
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